SCENTED LIVERWORT

SNAKE LIVERWORT [USA]

Conocephalum conicum

Liverworts [Conocephalaceae]

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5th Aug 2011, Cathedral Cave, Little Langdale, Cumbria. Photo: © RWD
Growing in a dark damp nook on limestone rocks in one of the entrances to this man-made cave, 'mined' from underneath for its limestone rocks (an upside-down quarry). (Hartstongue Fern growing in top corner for size comparison).


5th Aug 2011, Cathedral Cave, Little Langdale, Cumbria. Photo: © RWD
One of the largest liverworts, with thalli ('leaves') up to 17mm wide and often forms extensive mats. (Hartstongue Fern growing in top corner for size comparison).


5th Aug 2011, Cathedral Cave, Little Langdale, Cumbria. Photo: © RWD
Conspicuous thalli, much longer than broad, flat, leathery and dark green.


5th Aug 2011, Cathedral Cave, Little Langdale, Cumbria. Photo: © RWD
Thalli bifurcate (branch into two) at various intervals and have conspicuous 'pores' that make it look like a tongue or the skin of a snake. Older thalli tend to turn a muddy purple (top). The liverwort smells strongly aromatic, hence the common name.


5th Aug 2011, Cathedral Cave, Little Langdale, Cumbria. Photo: © RWD
Here growing with Hart's-Tongue Thyme-Moss.


5th Aug 2011, Cathedral Cave, Little Langdale, Cumbria. Photo: © RWD
The air pores are surrounded by a (mainly) hexagonal network of darker lines. Despite this, the surface feels smooth lacking texture.


5th Aug 2011, Cathedral Cave, Little Langdale, Cumbria. Photo: © RWD
Differs subtly from a very similar Liverwort, Conocephalum salebrosum, which was once lumped in with this one, see notes below.


5th Aug 2011, Cathedral Cave, Little Langdale, Cumbria. Photo: © RWD
Cathedral Cave from 'mine' adit. Warning notices advise of grave dangers within! Enter at your own risk and never alone!


Easily mistaken for : Conocephalum salebrosum and was in the recent past until in 2005 it was realised that there are two different species. [A third species, Conocephalum supradecompositum, if it exists, may not grow in the UK]. The differences between C. conicum and C. salebrosum are that conicum is somewhat larger, is shiny rather than matte when dry (its impossible to tell when wet), borders between air pores are less conspicuous, and that surface is smooth (rather than undulating where the 'valleys' between the pores are). C. salebrosum prefers slightly drier conditions than C. conicum. The British Bryological Society (BBS) list the differences on this page of their website.

The above specimens are believed by the Author to be those of C. conicum.

Both species are dioicous, with male and female organs being on separate plants.

This is a very common liverwort found almost throughout the UK, but tending to avoid the highest mountain areas of Scotland and Eire.

Habitat is moist or wet places that are in the shade most of the time, and on neutral or slightly basic substrates. Rocks and by rivers, waterfalls and streams, or damp banks or walls and on rocky ledges, boulders and in gullies.

ODOUR COMPOUNDS in SCENTED LIVERWORT

There exist three chemo-types of Conocephalum conicum, types 1, 2 and 3, which respectively emit Sabinene, Bornyl Acetate and Methyl Cinnamate as their major components, giving each type a differing characteristic odour when crushed. It is possible that each corresponds with the recent splitting into three differing species, C. conicum, C. salebrosum and C. supradecompositum but this is not made explicit in the literature. The odour is variously described as being camphoraceous, strongly mushroom and lactone-like, although which one corresponds to which chemical is similarly vague.

Sabinene is a bicyclic monoterpene consisting of a five-membered ring fused to a 3-membered ring and which is present in several plants such as Norway Spruce, Holm Oak, Tea Tree, and contributes to the spiciness of Black Pepper and Nutmeg. It is a major consistent of Carrot seed oil. It is used as a flavour and a perfume and has a woody pine-like fragrance with a spicy nuance.

Methyl Cinnamate is an ester of Cinnamic Acid and has a strong sweet fruity aromatic odour a little like that of Strawberry in which it also occurs, or Cinnamon. It is used in the perfume industry as a perfume and as a flavour, which is fruity again like Strawberry. Methyl Cinnamate also occurs in the Strawberry Gum Eucalyptus treem, in some varieties of Basil and in Sichuan Pepper.

It acts as a pheromone to male orchid bees.

Bornyl Acetate is another ester and has a pine-like camphoraceous odour. It is used in both perfumes and as a flavour.

ANY TEXT GOES HERE


Distribution
 family8Liverworts family8Conocephalaceae
BSBI maps
genus8Conocephalum
Conocephalum
(Marchanta Liverworts)

SCENTED LIVERWORT

SNAKE LIVERWORT [USA]

Conocephalum conicum

Liverworts [Conocephalaceae]

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